Satellite fly

This is a new one for me - like a lot of people I didn't even know such things existed.

Last month I was surveying a site in Surrey, just outside of Greater London, and had found a nice south facing slope with a thermophilic invertebrate community present. This included mining bees Andrena sp., plus the cleptoparasitic nomad bees Nomada sp. and bee-flies Bombylius sp., and the wasp Dolichovespula media. Also present were the ants Formica fusca, so I spent a bit of time on my knees seeing what other species were present (only Lasius niger s. str. and Myrmica scabrinodis as it turned out).

Whilst I was on the ground a solitary bee flew by, followed closely by this fairly nondescript looking fly. The bee stopped, and the fly stopped a few centimetres behind. The bee flew a little further and stopped, and the fly did likewise. It slowly dawned on me that the fly was actually stalking the bee!

The bee started to move down amongst some grass whilst the fly waited around, so I took the opportunity to take the photographs below. Okay, so they will win no prizes, but they were the only two I could take before I got too close and scared the fly away.

It's taken me a while to work out what this thing was, but I'm now pretty certain it was a satellite fly Leucophora sp. Like Nomada and Bombylius, Leucophora are cleptoparasitic, laying their eggs in the burrows of the bees they hunt. According to the Society for the study of flies there are only eight species of Leucophora in the UK, none of which have been commonly recorded if their maps are any judge (though these are likely to be very incomplete).

However, other than this I've been unable to find out any information. Remarkably, none of the British species of Leucophora have conservation status, which implies that they were missed or ignored the last time these flies were reviewed, as some of the eight species must be rare. Perhaps too little was known about them to make an accurate judgement of their status. Honestly, if it hadn't been behaving so unusually I'd have ignored this individual.

If anyone else has any useful information on these beasts I'd be very interested to hear, even if it is to say that they are actually really common!

Myopias tenuis

I've been pretty quiet recently as I've been very busy organising my first overseas anting trip of 2008. About a month ago, the University of Regensburg, Germany, invited me to visit them. Part of the trip will involve a seminar, so I've been trying to write the seminar, arrange travel, etc. and re-learn the little German I knew 15 years ago. This and my full-time job have left little time for very much else.

To make up for the lack of posting I decided to post something rather neat. The specimen in the picture is Myopias tenius. It wasn't collected by me; that honour goes to Catherine L. Parr, who also attended the Ant Course in 2007. Catherine found a number of interesting species during the two weeks and kindly let me have this specimen. I've only now got around to identifying it.

According the Shattuck (1999), Myopias are infrequently collected and are found from Sri Lanka to southeast Australia. Within Australia, M. tenuis is only known from north Queensland, but is also known from New Guinea, including the Indonesian part, and the Solomon Islands (Willey & Brown, 1983). This specimen was collected from Smithfield Conservation Park, an area of rainforest about 10 minutes walk from James Cook University, Cairns campus.

Roy Snelling

It is with sadness that I report the death of Roy Snelling. Roy was one of the great figures in the study of Hymenoptera. He died in his sleep at the start of an expedition in Kenya.

I only had a small amount of contact with Roy, so I won't say too much. I met him on the Ant Course in Cairns, 2006 and had some email contact with him afterwards. He was a brilliant man with a wicked sense of humour, so I liked him right away, as I expect most people did.

He was also very generous with his expertise: he resolved the identity of some problem Rhoptromyrmex from Queensland and also provided some useful, and sometimes entertaining discussion on North American Lasius. His initial reaction to my query was, 'I detest Lasius!', a sentiment I suspect most people can sympathise with!

It is a sad loss, all the more so for his family and those close to him, to whom my thoughts go out.

Eulogies are at Myrmecos Blog and The Ant Farm and Myrmecology Forum.

Blooo! Blong!

It must be my inner child. This video had me in tears.



I'm just not sure whether this counts as cruelty - my belly aches in sympathy.

Miscellaneous ant collections

The last of the boring but possibly useful lists.

This covers those counties where I only have a few species recorded or in my collection. Those from Ireland and Senegal were collected by me. My parents have been good enough to collect ants for me, including from Mallorca and Austria. Specimens collected by David M. King were passed to me for identification; these came from Italy, Kuwait, Macedonia, Qatar, Turkey and the Canary Isles (not yet included). Claes-Göran Magnusson sent me a specimen from Sweden. Finally, Sabine Frohschammer passed on a few specimens from a live colony she had collected in Malasia.

Austria

Formicinae
     Camponotus
          herculeanus
     Formica
          aquilonia
          cinerea
          exsecta
          lemani
          lugubris
          paralugubris
          sanguinea
     Lasius
          fuliginosus
          niger
          platythorax
Myrmicinae
     Leptothorax
          acervorum
     Manica
          rubida
     Myrmica
          lobulicornis
          rubra
          ruginodis
          scabrinodis
          sulcinodis
     Harpagoxenus
          sublaevis

Ireland

Formicinae
     Formica
          lemani
     Lasius
          flavus
          grandis ¹
          niger
Myrmicinae
     Myrmica
          ruginodis
          sabuleti
          scabrinodis
          schencki
     Tetramorium
          caespitum

Italy

Myrmicinae
     Aphaenogaster
          ionia

Kuwait

Myrmicinae
     Messor
          ebeninus
          minor
     Monomorium
          subcomae
     Pheidole
          teneriffana ²

Macedonia

Formicinae
     Cataglyphis
          aenescens

Malasia: Sarawak

Myrmicinae
     Monomorium
          floricola

Mallorca, Spain

Myrmicinae
     Crematogaster
          scutellaris
Dolichoderinae
     Linepithema
          humile ³

Qatar

Myrmicinae
     Monomorium
          tumaire

Senegal

Formicinae
     Camponotus
          sericeus
Myrmicinae
     Crematogaster
          senegalensis
Ponerinae
     Pachycondyla
          sennaarensis

Sweden

Formicinae
     Lasius
          umbratus

Turkey

Formicinae
     Cataglyphis
          nodus
Myrmicinae
     Aphaenogaster
          ionia
     Messor
          caducus
          denticulatus
     Pheidole
          pallidula
Dolichoderinae
     Tapinoma
          festae

---

¹ I know this seems unlikely, but it's the best fit. I've had a paper half written for two years now. My problem is that I can't make my head measurements match those of Seifert's - on any Lasius. I've spoken to others and this is not a problem restricted to me! Oh, and Ireland also has L. psammophilus, with no evidence for L. alienus.
² Introduced.
³ Introduced and apparently one of the dominant species on Mallorca now: my parents only collected the two species above, despite searching hard.

Australian ant collections

This is the most incomplete of the lists I'm going to present. There are two main reasons for this: the taxonomy of the Australian ant fauna is incredibly difficult, making identification hard, and I was interrupted by my work on the Gambian fauna. Eventually I will get back to these collections.

They are the result of my attendance on the Ant Course 2006, plus an extra week that I stayed. This was held at James Cook University (JCU) Cairns Campus. If you are interested in ants then I can't recommend the Ant Course strongly enough.

Perhaps not surprisingly, the Australian ant fauna is quite special. Groups that are rare elsewhere, such as Iridomyrmex and Polyrhachis have diversified into many different forms. Others, such as Leptomyrmex, Turneria, Melophorus and the famous bulldog ants Myrmecia are limited to the Australasian region. I should also make mention of possibly the most famous Australian ant, Nothomyrmecia macrops.

There is a lot of work still to be done on these specimens. Iridomyrmex, Camponotus, Polyrhachis, Crematogaster and Pheidole are all common and diverse in Australia, so I collected a lot of different species. A key now exists for Camponotus, but I've only just started trying to sort through all the specimens I have. Iridomyrmex in particular is well-nigh impossible, as there are many species that differ very little. Many of the smaller groups also lack keys, so identification is not easy.

For further coverage check Australian Ants Online.

I should say that the specimens actually belong to JCU and that I have them as part of a loan to Cardiff Museum. Once I've finished with them they will go to Cardiff or back to JCU.

Aenictinae
     Aenictus
          aratus
Cerapachyinae
     Cerapachys
Dolichoderinae
     Anonychomyrma
     Iridomyrmex
     Leptomyrmex
          unicolor
          varians subsp. ruficeps
     Ochetellus
     Tapinoma
          melanocephalum ¹
          minutum
     Technomyrmex
          albipes
     Turneria
          bidentata ²
Ectatomminae
     Rhytidoponera
Formicinae
     Acropyga
     Calomyrmex
     Camponotus
          aeneopilosus
          dromas
          vitreus
     Echinopla
          australis
     Melophorus
     Oecophylla
          smaragdina
     Opisthopsis
          haddoni
          jocosus
          pictus subsp. bimaculatus
     Paratrechina
     Polyrhachis
Myrmeciinae
     Myrmecia
Myrmicinae
     Cardiocondyla
          nuda ³
     Crematogaster
     Mayriella
          overbecki ⁴
     Meranoplus
          ajax
          hirsutus
     Monomorium
          fieldi
          laeve
          pharaonis ⁵
     Pheidole
     Podomyrma
     Rhoptromyrmex
          wroughtoni sp. 1
          wroughtoni sp. 2⁶
     Tetramorium
          deceptum
          lanuginosum ³
          simillimum ³
     Wasmannia
          auropunctata ⁷
Ponerinae
     Hypoponera
     Leptogenys
     Myopias
          tenuis
     Odontomachus
     Pachycondyla
Pseudomyrmecinae
     Tetraponera
          punctulata

---

¹ Introduced and collected in many situations, from wash-blocks to rainforest.
² I only collected one specimen of this rarity, which is usually found in the canopy. I found it on my clothes in the room I was staying in at (JCU) Cairns Campus. I suspect that someone had shaken the tree it was on as I was beneath.
³ Introduced.
⁴ A bit of a rarity, especially in Queensland, where it is otherwise unknown (Shattuck & Barnett, 2007).
⁵ Introduced, found once crawling across the desk in the room I was staying in at JCU.
⁶ Considered to be rare, but I found it frequently enough in the rainforest that by the time I left I could recognise it on sight. Following a chat with Barry Bolton (who last reviewed the genus), I feel confident in saying that what I have belongs to two species.
⁷ Introduced around Cairns and apparently presenting quite a serious threat. Found abundantly in one place near JCU.

French ant collections

This list of ants in my collection was the result of a two week 'holiday' in the Dordogne and Massif Central areas of France. It was the first time I had ever collected ants outside of the UK and forecast the end of 'normal' holidays for me!

More information on the ants of France can be found at Fourmis, the French ant forum.

Dolichoderinae
     Dolichoderus
          quadripunctatus
     Tapinoma
          erraticum
Formicinae
     Camponotus
          aethiops
          cruentatus
          fallax
          gestroi ¹
          lateralis
          ligniperda
          piceus
          trunctatus
          vagus
     Cataglyphis
          cursor
     Formica
          fusca
          gagates
          pratensis
          rufa
          rufibarbis
     Lasius
          alienus
          brunneus
          emaginatus
          flavus
          fuliginosus
          grandis ²
          niger
          platythorax
          psammophilus
     Plagiolepis
          pygmaea
Myrmicinae
     Crematogaster
          scutellaris
     Messor
          structor
     Myrmica
          sabuleti
          scabrinodis
     Pheidole
          megacephala ³
          pallidula
     Temnothorax
          grouvellei
          nylanderi
          racovitzai
          unifasciatus
     Tetramorium
          caespitum

---

¹ Two alate queens, one found in the Salignac-Eyvigues, Dordogne, the other on Causse Noir in the Massif Central. The latter was found in association with C. lateralis. This species is not supposed to occur in mainland France, but after a lot of debate they were eventually confirmed by Cedric Collingwood.
² A rarely recorded species in France. This collection was from the edge of a woodland near Nant, Massif Central. This has also been confirmed by Cedric Collingwood and matches the morphometrics given in Seifert (1992).
³ An introduced species in France, found alongside P. pallidula on the château walls at Castelnaud, Dordogne.