Monomorium dakarense

I previously thought I had collected Monomorium dictator in the Gambia, as that is what this specimen keyed out as. However, I've been doing a lot of work on the salomonis-group recently and I no longer think that I was correct. I now think it is Monomorium dakarense.

It all boils down to couplet 36 of Bolton's (1987) key. I think it would be possible to debate for hours over whether the sculpture on the head of this specimen is 'uniformly densely reticulate-punctate' or 'with a silky, smeared or roughened appearance'. Initially I opted for the former because, well, it is reticulate-punctate, just more finely than the likes of Monomorium bicolor! However, I've since come to realise that this is what Bolton means by the latter, which leads to M. dakarense.

The difference between M. dakarense and M. dictator, other than the ambiguities of cephalic sculpture, seems to be size, number of ommatidia in the longest row and, importantly, scape index (SI). M. dakarense is overall the smaller of the two, with fewer ommatidia and a SI of 95-100, whereas M. dictator has a SI of 107-109. For those not familiar with SI, it is as follows:

Scape length × 100

Head width

The SI in this specimen is 92, putting it closest to the range given for M. dakarense. Given that Bolton only had the three syntype workers to measure it is likely that the true range of the SI includes 92. However, this is not the only difference between this specimen and the holotype, as the other measurements are (quoting the measurements for MJL143, followed by the range given by Bolton, in mm):

     Head length = 0.61 (0.57-0.59)
     Head width = 0.51 (0.44-0.47)
     Cephalic index (ratio of head width to height) = 84 (77-80)
     Scape length = 0.47 (0.44-0.45)
     Pronotal width = 0.31 (0.30-0.31)
     Mesosoma length = 0.72 (0.66-0.70)

Despite the larger size, the specimen is not big enough to be M. dictator and differs in the other features mentioned, so it is most likely M. dakarense.

It still remains a bit of a rediscovery. M. dakarense had been collected once in Senegal in 1914, so it's more likely than M. dictator, which had only been collected in Angola. The only specimens that I collected were found on the ground at Madiyana Camp on Jinack Island, and only at night. Since we stayed at Madiyana Camp for a few days it would be surprising if M. dakarense had been active during the day and I had not collected it then, which indicates pretty strongly that it is entirely nocturnal. Perhaps a combination of its nocturnal habits and its small size have meant that it has not been found again until now?

Monomorium sp. A

When I first attempted to identify this ant I decided that it was Monomorium exiguum, though I wasn't exactly convinced. I was using Bolton (1987), though I find the key fairly vague at times, which isn't helped by the fact that the descriptions aren't exactly consistent and many species lack diagrams.

M. exiguum is the last species in the key - okay, so there aren't that many Afrotropical Monomorium with 11 antennal segments, but even so I feel that if you've got to the end of the key and your not convinced, then chances are you are wrong.

Then I discovered that I had collected specimens that were a perfect match for M. exiguum and very different from this species, so it was clear that I was wrong. I went back to the key and decided that the next best fit was Monomorium dolatu, though I was rather less than convinced - the petiole shape and Cephalic Index appeared to be correct, but there were slight differences in the structure of the clypeus.

My trip to the Natural History Museum, London has thrown the latest spanner in the works, as I've now been able to examine specimens that Bolton identified as M. dolatu closely. It's pretty clear that this isn't M. dolatu, as the petiole shape is not near as conical as on that species and it's generally wrong. I was also able to check and eliminate M. pulchrum, M. taedium, M. fastidium and M. vaguum. I have in my collection and was able to confirm M. rosae, M. exiguum and M. mictilis. The only other species that it could remotely be is M. bequaerti, which was not present in the museum collection, but this should have a postpetiole the same size as the petiole, as in M. pulchrum, which my specimen does not have.

Since I've eliminated all Afrotropical species of Monomorium with 11 antennal segments I'm having to call it 'species A' for the moment, at least until I've checked those species occurring elsewhere. This is the second potentially new species of Monomorium I've encountered in as many months - the first I've already established is definitely new.

M. sp. A was collected once, nesting within an Acacia pod, in Kololi. Because this was within our last few hours in Gambia and I had packed everything to leave, they were collected alive. I had to carry the pod, with the ants, back to the hotel in my hand. This involved dodging the bumsters (notorious in The Gambia) and trying to keep the pod safe whilst I had a chat with a charming young Gambian woman, who invited me to a 'club' and I suspected to be a prostitute. Once I made it back to my room I put the pod and the ants in a bag and in my case, where they remained until I got home.

I kept them alive for a few days before realising all I had were a few workers - not worth the effort of keeping long-term. They were also a pain; they proved to have an incredible ability to walk upside down on a smooth surface and vertically across fluon. This, combined with the fact that they tended to wedge themselves into the small gap between container and lid made them very difficult to contain. Given these recorded habits it seems that this species is well adapted to an arboreal lifestyle.

Incidentally, the droplets of liquid are grease. I will have to do something about this soon, whereupon I may upload some fresh photographs.

Camponotus vestitus subsp. intuens

I originally identified this as Camponotus cosmicus and wrote a nice long blog defending my decision to call it that. I even provisionally synonomised it with a few other species and subspecies. I was wrong!

At the time, I sent photographs to Brian Taylor, who replied, 'Why do you not think this is Camponotus vestitus?' Perhaps I should have recognised that his far greater experience was likely to lead to a correct identification, but I like to learn from my own mistakes.

C. cosmicus was one of the species that I checked when I visited the London Natural History Museum, as the holotype is there. C. cosmicus is a large and fairly stocky ant, even the minors, unlike my specimens. I was also able to check C. vestitus subsp. pectitus, which has all the right features (divergent pubescence on the gaster, same number of setae on the mesosoma, same propodeum shape) and differs only slightly in the colour of the head.

Arnold (1924) stated that Smith's (1858) description of C. cosmicus is useless, and it appears as though he is right. However, none of the authors who have written about C. cosmicus picked up on what I believe is probably the most significant characteristic, as all specimens in the museum had many erect setae on the ventral surface of the head. I can also add that C. cosmicus is predominantly very dark red (not black), with the propodeum rounding smoothly from the metanotal groove, less divergent pubescence on the gaster and a greater number of setae on the mesosoma.

So, maybe it is C. vestitus subsp. pectitus. It gets complicated because Bolton (1995) states that Santschi (1926) suggested that C. vestitus subsp. pectitus and C. vestitus subsp. intuens are the same, but at no point do they appear to have been properly synonymised. Reading Santschi's statement, it seems pretty clear that he has no doubt, so I can only assume that Bolton did not synonymise it because the description of C. vestitus subsp. pectitus post dates this statement! Forel's (1909) description of C. vestitus subsp. intuens is just as ambiguous as the descriptions of C. cosmicus and C. vestitus subsp. pectitus.

Update Nov 2008: My specimens have now been compared with the type specimens from the Naturhistorisches Museum Basel. This allowed me to confirm that they are very clearly C. vestitus subsp. intuens - a completely identical match. Possibly the colour of the head is the key feature in discriminating these subspecies.

I also showed my specimens to Cedric Collingwood, who claimed them to be C. jizani. This is a species from the Middle East, though he admitted that the only reason he named it thus was because he didn't know what else to call it! Possibly, C. jizani is a junior synonym of C. vestitus subsp. intuens or C. vestitus.

This species was collected a number of times only in the grounds of the hotel we were staying at in Kololi, Gambia. Here it could be found nesting in turret nests, so named because there were steep 'craters' of sand piled around the nest entrances, up to about an inch high. These are described by Arnold (op. cit.) for C. vestitus subsp. pectitus, which he states is rare and found nesting in sandy soils. He then describes,

'nest-entrances[s] surrounded by a circular, high and sharp-edged crater... This form of crater is distinctive of this insect.'

Corrections

Before I disappeared for Easter I visited the London Natural History Museum to compare my some of my Gambian specimens with ant specimens in their collection. As a result, I have a few changes to make, but need to do a bit more work before I make them. Until I change the posts, this will hopefully caution against relying on previous posts.

I had misidentified the following species:

• Camponotus cosmicus - more likely within what can best be described as the vestitus complex.
• Monomorium dolatu - doesn't appear to be any of the species included in Bolton (1987). I had suspected that this was the case, but without checking reference material went with the closest option.

I had correctly identified these species:

• Monomorium exiguum
• M. rosae
• M. osiridis
• M. vonatu
• M. egens
• Dorylus affinis

These species are perhaps open to debate, as I couldn't conclusively establish whether they were the same species or different:

• Camponotus rufoglaucus controversus - slight differences in setae and pubescence but the specimens checked were not types and were majors only.
• Camponotus olivieri lemma and delagoensis - I couldn't check these subspecies.
• Crematogaster impressa - the specimens present were collected by Donisthorpe and did not appear to be the same as mine or each other. Chances are this species and C. excisa will not be resolved until the taxonomy is revised.

Camponotus olivieri

Camponotus olivieri is a widespread species in Africa, ranging from South Africa north to Sudan and west to Ghana. However, it is made up of a great many subspecies that probably deserve species status, so the situation may be rather more complicated than it seems at first.

Two subspecies of C. olivieri were collected in The Gambia: lemma and delagoensis. They differ quite markedly in pubescence and also, to a lesser extent, morphology. What I think provides some evidence that they deserve separate status is that they were both found at one site.

C. olivieri subsp. lemma was found in two locations: Abuko National Park, Lamin and Bijilo Forest Park, Kololi. In both cases it was found only on the ground. It can be distinguished from other species of C. olivieri by the rust colour at the base of the antennal scapes and the gaster, which is uniformly shiny, sparsely pubescent but still more densely pubescent than the mesosoma and has erect hairs that are thicker than the pubescence.

C. olivieri subsp. delagoensis was found at Abuko National Park, on a tree. It is similar to subsp. lemma, but has denser, yellow pubescence, especially on the gaster.

Both ants have been recorded only rarely, possibly due to the fact that they are regarded as subspecies and so have been more frequently recorded at species level. C. olivieri subsp. lemma had been recorded in Angola, Tanzania and former Zaire. C. olivieri subsp. delagoensis was only known from the type location at Delagoa Bay in Mozambique. This makes these Gambian records an extension of the known range of the species and both subspecies, especially large in the case of subsp. delagoensis.

These have both been compared with the type specimens from the Muséum d'histoire naturelle, Genève and the Naturhistorisches Museum Basel. This allowed me to confirm C. olivieri subsp. delagoenisis.

The syntypes of C. olivieri subsp. lemma are rather variably hairy, so much so that I wondered if Forel had mounted specimens from different colonies on the same pins. The main difference between these and my specimens was that most of the types were much less hairy, though one was similar to mine. It could be that the specimens have lost some of their pilosity. Regardless, I feel I need to do some more work to confirm that there is nothing closer.

Plagiolepis mediorufa

Plagiolepis mediorufa had only been recorded from the Congo and the Central African Republic, generally from myrmecophilous plants. My Gambian specimens were collected from plants, trees and a palm frond in Bijilo Forest Park, Kololi and Abuko National Park, Lamin, but nothing about the plants indicated that they were adapted to supporting ants. The Gambia also represents a significant range extension north and westwards.

The trouble is that I suspect there may be a few, as yet undescribed, small yellow and morphologically similar species of Plagiolepis in Africa as a whole. I have placed my specimens in P. mediorufa because it is currently the best fit, but would be interested to see where they are placed if the genus is ever reviewed.

The specimen that I photographed was sadly rather battered. Plagiolepis often tend to be rather soft and fragile, and so shrivel when they dry. Tapinoma does the same thing. (I think the funiculus of the antenna was missing when I collected it.) It was still the best of the three specimens that I had mounted.

Monomorium bicolor

Monomorium bicolor is an abundant ant in The Gambia. It had been collected there before in around 1926 in Banjul (formerly Bathurst), and in 2007 in Kololi and on Jinack Island. It is distributed from Angola north to Sudan and from Somalia across to Senegal.

M. bicolor seems to be present in a variety of habitats, as it was found on trees, on the ground in open areas, on buildings and once scavenging from a dead fish, but was only found nesting in the ground.

M. bicolor is very similar to Monomorium dictator, but differs by being generally larger, though also variable in size, and lacking abundant setae on the anterior half of the first gastral tergite. It should have at least two erect setae on the anterior margin of the first gastral tergite, but these were lacking in all Gambian specimens. Five out of the nine specimens examined had distinct pits where the setae should have been, but all lacked setae. Nevertheless, there is no doubt that they are M. bicolor, so the hairs have perhaps rubbed off or Bolton (1987) defined the species too narrowly.

Monomorium osiridis

Monomorium osiridis had only been collected twice, in Kenya, before I picked up these specimens in The Gambia. It seems quite surprising to find M. osiridis on the other side of the continent, but I don't know what else these specimens could be.

M. osiridis and the southern African species Monomorium zulu and Monomorium rabirium are separated from other Monomorium by having 12-segmented antennae, smooth mandibles, conspicuous eyes, sculptured propodeum and no standing hairs on the dorsal mesosoma and gaster. M. osiridis differs from the other two by having a sculptured head, except for a median strip (visible in the bottom photograph).

I only collected M. osiridis once, on the ground at Madiyana Camp on Jinack Island. They were collected only at night, strongly indicating that they are entirely nocturnal, as may be the case with Monomorium dictator. This could be another instance of a species being under-recorded partly because it is nocturnal.

These have been compared with specimens checked by Bolton at the Natural History Museum, London, and are confirmed as M. osiridis.

Monomorium rosae

Monomorium rosae is a widely distributed species with a central African distribution that stretches from Kenya to Senegal. It was collected twice on Jinack Island in The Gambia, once on a tree and once in dry leaf litter at the base of a tree.

Bolton (1987) notes that this species varies in size and pilosity over its range, suggesting that M. rosae may contain more than one species. He distinguishes the species as currently defined as having 11-segmented antennae, dark colour, moderately long scapes and a distinctively shaped postpetiole.

It's interesting how many of the Monomorium now known from The Gambia have 11-segmented antennae. M. dolatu, M. exiguum, M. mictilis and M. rosae all have 11-segmented antennae, out of ten species recorded from The Gambia (40%). This contrasts with 12 species out of 149 listed by Bolton for Africa as a whole (8%). Maybe Monomorium with 11-segmented antennae are more common in this part of west Africa?

These have been compared with specimens checked by Bolton at the Natural History Museum, London, and are confirmed as M. rosae.

Cataglyphis oasium

The size of an ant and how easy it is to identify are not necessarily related. Whilst Monomorium mictile, at just over 1 mm, was relatively easy to identify, I'd been pondering this very large species of Cataglyphis for weeks.

Cataglyphis is a genus that is taxonomically muddled. I've finally concluded that this species is probably Cataglyphis oasium. It was the keys in Santschi (1929) and Radchenko (1998) that eventually provided the most likely identification.

C. oasium was originally described by Santschi (1929), though the name he gave it (Cataglyphis bicolor st. nodus v. oasium) was invalid, so the authority for this species is Menozzi, 1932. Menozzi only gave two lines of text to the species, changing the name to Cataglyphis bicolor var. oasium, without noting that it was a new combination or providing a fresh description, and naming Santschi as the authority. The variety oasium was later given subspecies status and then species status by Radchenko (1997). Wehner, Wehner & Agosti (1994) suggested that C. bicolor subsp. oasium was synonymous with Cataglyphis savignyi, though they didn't actually synonomise the two names. Thus, the valid name is C. oasium. It's very confusing, so it took a little while just to sort out this history.

Information on the distribution of C. oasium is very sparse. Santschi mentions records from Tunisia and Algeria, whilst the most recent record that I'm aware of is Libya in 1931, which is reported by Menozzi. Radchenko (1998) states that it is found in North Africa and the Middle East.

This gives the impression that C. oasium is rare, but this seems unlikely. It is probably under-recorded due to the taxonomic difficulties. It is possible that the species collected by Lenoir and mentioned in The Ants of Africa is also this species. If C. oasium is found in oasis in the Sahara and in the Middle East it seems reasonable that it could also occur in The Gambia and Burkina Faso.

The Gambian specimens were collected from Kololi and Madiyana Camp on Jinack Island, mainly from sandy ground in scrub savannah. In this sort of hot scrub savannah it could be quite common, so it may be abundant further inland, away from the coastal breeze.

Extreme myrmecology: Monomorium mictile

Monomorium mictile has to be the smallest ant I have ever found. It is very inconspicuous.

I started off with four specimens, but misplaced two before I could look at them properly (I still don't know how). I selected one of the two specimens to be mounted and, whilst I was getting everything ready, I brushed a bit of dust off my microscope stage. I knew what I had done as soon as I had done it - the bit of dust was the ant. It was no good looking for it on the floor, as it was gone.

This left me with just one, which had not died in a position that was conducive to mounting and I felt that trying to fix this would be too risky on such a small specimen. This is why the specimen in the photograph is not as well mounted as most others.

Bolton (1987) comments that material of this species is relatively scarce; it's not hard to see why!

M. mictile is widespread in Africa, found from South Africa north to Mali and Sudan. It as collected once in The Gambia from a tree in the coastal scrub on Jinack Island.

Monomorium exiguum

Monomorium exiguum has been found across central and southern Africa, from Ethiopia in the east to Guinea in the west and Zimbabwe in the south. It appears to be primarily a forest species, so it is no surprise that the Gambian specimens were collected from a tree in Abuko National Park, one of the few remaining patches of gallery forest.

Bolton (1987) states that M. exiguum as defined definitely contains more than one valid species, but could not resolve them. It is therefore interesting to note the small size of the Gambian specimens, which are apparently much smaller than is typical for the species; Bolton gives lengths of 1.5-1.7 mm whereas the Gambian specimens can be little more than 1 mm long.

These have been compared with specimens checked by Bolton at the Natural History Museum, London, and are confirmed as M. exiguum.

Monomorium vonatu

Yet another rediscovery. It gets too familiar after a while, but it does demonstrate how little is known about the ants of Africa.

Monomorium vonatu was known only from a single holotype specimen in the Natural History Museum, London. It was collected in 1970 in Ghana and described by Bolton (1987). It is easily distinguished from other species of Monomorium by the shape of the petiole and postpetiole, which both have distinct ridges running across them. The only other species that has this structure is Monomorium mirandum, which is very distinctly bicoloured.

As if to demonstrate that you don't need to go to great lengths to find interesting species, I collected my M. vonatu specimens in the grounds of the hotel we were staying in. They were found on the ground, either in bare sand or grassy areas.

Quite why this species hasn't been discovered anywhere else seems a bit of a mystery, as it was probably pretty common in suburban Kololi. Perhaps it is overlooked as another small black species of Monomorium.

These have been compared the holotype at the Natural History Museum, London, and are confirmed as M. vonatu.

Slightly confused Monomorium egens

This specimen has proved difficult to identify, but I'm now pretty certain that I've got it right. It keys out pretty clearly as Monomorium egens using Bolton (1987), but is generally much more slender than the species that Bolton describes and has much longer scapes (SI 108).

Because it doesn't fit perfectly I've been puzzling over it for the past couple days. The resolution came when I translated Santschi's (1926) description of M. longiusculum from the French. This describes a slender species with scapes long enough to reach the posterior margin of the head (I've included my rough translation of Santschi's description below). This means that my specimen has slightly longer antennal scapes and may be more slender, but is probably close enough.

Bolton synonomised M. longiusculum with M. egens. What is strange is that he doesn't mention this level of variety despite having examined the type material for M. longiusculum, though he does confess not to be convinced that M. egens is a single species. Having now compared my specimens with those checked by Bolton in the Natural History Museum, London, I can understand why he decided to synonymise them, as there is no obvious difference other than the shape.

This species is another forest species from Abuko National Park in The Gambia, where it was collected from a tree. Bolton states that M. egens nests in rotten wood in the soil or in fallen trunks and forages in the wood and leaf litter. This seems to make finding it part way up a living tree trunk quite usual.

M. egens has been found in West Africa from Angola to Guinea, putting this specimen at the extreme northern edge of its known range.

The following is my translation (with a little help from Brian Taylor) of the type description for M. longiusculum from the French.

Monomoriumlongiusculum Santschi 1926

Worker. Length: 2.1 mm. Body brown, gaster brown-black. Appendages yellowish-brown. Funiculus yellowish. Smooth, shining. Slightly pilous. Pubescence suberect and more abundant on the scapes, adpressed and sparse on the legs.

Head approximately a quarter longer than broad, rather convex at the sides, the posterior margin straight with the angles rounded. Eyes two thirds as large as the space which separates them from the anterior margin of the head and placed between the middle and a third back from the anterior margin. Carina on the clypeus marked, not very divergent and little or not projecting at the anterior margin. Mandibles with 4 teeth. The scape reaches the posterior margin of the head. Articles 2 to 9 of the funiculus about as thick as long. Pronotum 'sub-shouldered', without a promesonotal suture and with the mesonotum in profile forming a regular and rather low convexity from the anterior to the metanotal groove. Propodeum longer than two thirds of the promesonotum and low, also convex, the sloping face not very distinct from the declivitous one; convex from one side to the other and narrow, width about half its length. Peduncle of the petiole as long as half of the base of the node; node triangular in profile and longer than high, not convex ventrally. Postpetiole a little shorter and lower than the petiole, roughly as high as long and rounded at the dorsally, as broad as long and a little broader than the petiole.

Monomorium afrum

Monomorium afrum is the largest species in its genus found in the Gambia so far. It is very widespread in Africa, being from from South Africa to Sudan. The closest it has been found to The Gambia is the Ivory Coast, so this find represents an extension of its known range westwards.

It was found only once as a worker, running on the ground in the coastal resort of Kololi. Other species of Monomorium are much more common in The Gambia.

The sculpturation of the head and mesosoma is interesting, as it makes even in focus pictures appear out of focus. This reticulate-punctate sculpturation can be seen on the close up photograph of the head.

On the same day and in the same location as the M. afrum worker was found I collected a lone dealate queen. Queens away from colonies present problems for identification, as most have not been described. For this reason, I left the Monomorium queens that I collected until last (within the Monomorium at least). Fortunately, the queen of M. afrum has been described by Arnold (1926).

What's more, it is incredibly distinctive. In fact, when I first looked at it, I assumed that it was a Tetramorium, at least until I realised that it had the single unpaired seta projecting from the midpoint of the anterior clypeus typical of Monomorium and lacked the raised clypeal ridges of Tetramorium. It is the most peculiar Monomoroium that I have ever encountered.

My specimen seems to differ slightly from those described by Arnold. He describes the second and third abdominal segments of M. afrum queens being anteriorly smooth and shining. This presents a problem, as the second and third abdominal segment is the petiole and postpetiole. However, authors are sometimes unclear about this, so Arnold could have been referring to the second and third gastral segments (abdominal segments five and six). Despite this uncertainty, I can find no smooth and shining areas on any part of my specimen.

The queen of M. afrum is obviously very different from the worker. Arnold states that had he collected a specimen without the context of workers and other queens he would have considered it to be the type of an entirely new genus, and it's easy to understand why. Bolton (1987) comments that it shows modifications characteristic of socially parasitic species of Monomorium and may prove to be a temporary social parasite.

Pachycondyla sennaarensis

Pachycondyla sennaarensis is probably one of the most common ant species in the coastal region of The Gambia, as I collected it 10 times. I also got to know it over the week that I was there and had soon started to ignore the many lone workers that I frequently saw.

Remarkably, given how common it is in the area that other myrmecologists were most likely to visit, it hadn't been recorded from The Gambia before. I'd be tempted to suggest that it might have spread into the country since the early 20th century, when the last myrmecologists visited, but it was recorded in Senegal at about that time. Perhaps it's just been more successful than other ants as the area has been developed.

It seemed equally happy around buildings and in natural areas, where it nested directly in the ground, often in cracks in concrete. It was described by Dejean & Lachaud (1994) as being at least partially seed eating, which is extremely unusual for a ponerine ant. My observations agree with this, as the refuse piles around nest entrances often contained seed husks and other vegetable matter.

They also showed the kind of wear on their teeth that other seed eaters, such as Messor, tend to develop on older workers. Rather typically, it proved impossible to get a decent photograph of the one specimen that showed really distinctive wear. I hope that the photos below give some indication of what I mean, despite the fact that they both chose to die chewing on their own legs.

The first one shows little wear:

Whilst the second shows slightly more (note particularly the tooth on the right just above the leg):

Perhaps you'll just have to take my word for it.

I also found a queen. This was washed up on the shore on the part of Jinack Island that is within Senegal, though it probably washed down the river Gambia to get there, as it was already dead. Despite the fact that it was not found in a colony, it has been possible to identify it as P. sennaarensis with some confidence, as none of the defining features are on the mesosoma and so are presumably the same on the queen as the worker.

This species also stings. I discovered this on my first day in The Gambia when, in my rush to get out and explore, I left my forceps and pooter in my case in the hotel. Picking up these with your fingers is perhaps not to be recommended! In terms pain, it wasn't actually too bad initially (I've been stung by worse), perhaps rating as a 2.5 on the Schmitt Sting Pain Index (just above a European honey bee). However, after 20 or so my thumb, in particular, was starting to feel a bit numb, so I decided to stop. For the next 3 days the skin on my thumb and forefinger, which had taken the bulk of the punishment, slowly peeled away.

All good fun.

Nesomyrmex angulatus

Nesomyrmex angulatus is another widespread species, found from South Africa to Saudi Arabia. The closest it has previously been found to The Gambia is Ghana, so this represents another western range extension (along with Cataulacus traegaordhi). The key thing about this species that separates it from other Nesomyrmex is the very smooth profile of the mesosoma, coupled with the yellow colour.

Though I only found it twice, it's probably fairly common in more wooded parts of The Gambia. It tends to be pretty inconspicuous. I collected it from trees in Bijilo Forest Park and on Jinack Island, from coastal forest and scrub savannah respectively.

Tetramorium sericeiventre

Tetramorium sericeiventre is a very widespread species, occurring from South Africa north to Mali. A single worker had been collected in The Gambia in 1926, recorded as subsp. arenarium.

This species is not as common as some ants in The Gambia, but can be found in Kololi and on Jinack Island, where it can be quite conspicuous. It was collected from savannah, where it nested in the ground.

I was lucky enough to retrieve a queen from one of the nests, which is shown in one of the photographs.

Cataulacus traegaordhi

This is the second Cataulacus collected from Abuko National Park in The Gambia (the other one was C. guineensis). I only managed to find one specimen of C. traegaordhi, so either it's less conspicuous, rarer or I just wasn't looking in the right places.

It has been found from South Africa north to Sudan, but apparently the closest it has been found to The Gambia is Ghana. Whilst finding C. guineensis in The Gambia apparently represented an northward extension of its known range, finding C. traegaordhi appears to have extended its known range westward.

Cataulacus guineensis

At the risk of unintentionally starting a theme that could potentially be very restricting and uninteresting, I think I'm going to continue to post 'ant portraits'. Someone might actually find them useful.

This one is Cataulacus guineensis. It was collected Abuko National Park, one of the more famous protected areas in The Gambia. Abuko is a remnant of gallery forest, rather than rainforest. Despite its small size and the fact that we hired a guide (which for me meant that I didn't really have enough time to stop and collect ants), I still managed to pick up some species of interest.

Three of these, C. guineensis, Cataulacus traegaordhi and Pyramica maynei are proper forest species, so finding them in one of the few remaining pieces of proper forest in The Gambia was quite satisfying.

C. guineensis has been found in the Republic of Guinea, as might be expected from its name, and throughout west Africa, but not from Senegal or further north, so this find from The Gambia may represent the northern limits of its known range.